B. W. Abbott, J. B. Jones, E. A. Schuur, F. S. Chapin, . Iii et al., Biomass offsets little or none of permafrost carbon release from soils, streams, and wildfire: An expert assessment, Environmental Research Letters, vol.11, 2016.

A. A. Ali, C. Carcaillet, J. L. Guendon, Y. Quinif, P. Roiron et al., The early Holocene treeline in the southern French Alps: New evidence from travertine formation, Global Ecology and Biogeography, vol.12, pp.411-419, 2003.

J. E. Andrews, Palaeoclimatic records from stable isotopes in riverine tufas: Synthesis and review, vol.75, pp.85-104, 2006.

N. B-elanger, C. Carcaillet, G. A. Padbury, A. N. Harvey-schafer, and K. J. Van-rees, Periglacial fires and trees in a continental setting of central Canada, Upper Pleistocene, Geobiology, vol.12, pp.109-118, 2014.

A. Berger and M. F. Loutre, Insolation values for the climate of the last 10 million years, Quaternary Science Reviews, vol.10, pp.297-317, 1991.

H. H. Birks and H. J. Birks, Future uses of pollen analysis must include plant macrofossils, Journal of Biogeography, vol.27, pp.31-35, 2000.

H. J. Birks and K. J. Willis, Alpines, trees, and refugia in Europe, Plant Ecology and Diversity, vol.1, pp.147-160, 2008.

O. Blarquez, C. Carcaillet, B. Mourier, L. Bremond, and O. Radakovitch, Trees in the subalpine belt since 11 700 cal. BP: Origin, expansion and alteration of the modern forest. The Holocene, vol.20, pp.139-146, 2010.
URL : https://hal.archives-ouvertes.fr/hal-01061557

L. D. Brubaker, P. M. Anderson, M. E. Edwards, and A. V. Lozhkin, Beringia as a glacial refugium for boreal trees and shrubs: New perspectives from mapped pollen data, Journal of Biogeography, vol.32, pp.833-848, 2005.

J. F. Buoncristiani and M. Campy, The palaeogeography of the last two glacial episodes in France: The Alps and Jura, Quaternary glaciations extent and chronology, pp.80059-80068, 2004.

R. Cantonati, S. Segadelli, K. Ogata, H. Tran, D. Sanders et al., A global review on ambient Limestone-Precipitating Springs (LPS): Hydrogeological setting, ecology, and conservation, Science of the Total Environment, vol.568, pp.624-637, 2016.

C. Carcaillet, A. A. Ali, O. Blarquez, A. Genries, B. Mourier et al., Spatial variability of fire history in subalpine forests: From natural to cultural regimes, vol.16, pp.1-12, 2009.
URL : https://hal.archives-ouvertes.fr/hal-01982876

C. Carcaillet and O. Blarquez, Fire ecology of a tree glacial refu, New Phytologist, vol.216, pp.1281-1290, 2017.

C. Carcaillet, J. L. Vernet, and . Willis, Comments on "The full-glacial forests of central and southeastern Europe, Quaternary Research, vol.55, pp.385-387, 2000.

P. U. Clark, A. S. Dyke, J. D. Shakun, A. E. Carlson, J. Clark et al., The last glacial maximum, Science, vol.325, pp.710-714, 2009.
URL : https://hal.archives-ouvertes.fr/hal-01845766

E. Cossart, M. Fort, D. Bourles, R. Braucher, and L. Siame, Deglaciation pattern during the Lateglacial/Holocene transition in the southern French Alps. Chronological data and geographical reconstruction from the Clar ee Valley (upper Durance catchment, southeastern France), Palaeogeography Palaeoclimatology Palaeoecology, pp.109-123, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01196633

E. Dahl, The nunatak theory reconsidered, Ecology Bulletin, vol.38, pp.77-94, 1987.

B. A. Davis, S. Brewer, A. C. Stevenson, and J. Guiot, The temperature of Europe during the Holocene reconstructed from pollen data, Quaternary Science Reviews, vol.22, pp.1701-1716, 2003.

M. E. Edwards, W. S. Armbruster, and S. E. Elias, Constraints on post-glacial boreal tree expansion out of far-northern refugia, Global Ecology and Biogeography, vol.23, pp.1198-1208, 2014.

J. Eikenberg, G. Vezzu, I. Zumsteg, S. Bajo, M. Ruethi et al., Precise two chronometer dating of Pleistocene travertine: The 230Th/234U and 226Raex/226Ra(0) approach, Quaternary Science Reviews, vol.20, issue.01, pp.20-28, 2001.

N. Fauvart, A. A. Ali, J. F. Terral, P. Roiron, O. Blarquez et al., Holocene upper tree-limits of Pinus section sylvestris in the Western Alps as evidenced from travertine archives, Review of Palaeobotany and Palynology, vol.169, pp.96-102, 2012.

P. Fliche, Flore des tufs du Lautaret (Hautes-Alpes) et d'Entraigues, vol.9, pp.387-400, 1904.

D. G. Gavin, M. C. Fitzpatrick, P. F. Gugger, K. D. Heath, F. Rodriguezsanchez et al.,

J. W. Williams, Climate refugia: Joint inference from fossil records, species distribution models and phylogeography, New Phytologist, vol.204, pp.37-54, 2014.

B. Van-geel, A. Protopopov, V. Protopopova, I. Pavlov, J. Van-der-plicht et al., Larix during the mid-pleniglacial (Greenland interstadial 8) on Kotelny Island, northern Siberia, Boreas, vol.46, pp.338-345, 2017.

B. Ghaleb and C. Falgu-eres, Apport des m ethodes bas ees sur le d es equilibre radioactif ( 238 U-234 U-230 Th-226 Ra-210 Pb) aux etudes des variations et changements climatiques, Anthropologie, vol.121, pp.73-81, 2017.

E. Gobet, W. Tinner, C. Bigler, P. A. Hochuli, and B. Ammann, Early-Holocene afforestation processes in the lower subalpine belt of the Central Swiss Alps as inferred from macrofossil and pollen records. The Holocene, vol.15, pp.672-686, 2005.

M. Hallinger, M. Manthey, and M. Wilmking, Establishing a missing link: Warm summers and winter snow cover promote shrub expansion into alpine tundra in Scandinavia, New Phytologist, vol.186, pp.890-899, 2010.

A. Hampe, F. Rodriguez-sanchez, S. Dobrowski, F. S. Hu, and D. G. Gavin, Climate refugia: From the last glacial maximum to the twenty-first century, New Phytologist, vol.197, pp.16-18, 2013.

O. Heiri, W. Tinner, and A. F. Lotter, Evidence for cooler European summers during periods of changing meltwater flux to the North Atlantic, Proceedings of the National Academy of Sciences of the United States of America, vol.101, pp.15285-15288, 2004.

A. Inderm?-uhle, T. F. Stocker, F. Joos, H. Fisher, H. J. Smith et al., Holocene carbon-cycle dynamics based on CO 2 trapped in ice at Taylor Dome, Antarctica, Nature, vol.398, pp.121-126, 1999.

L. Ju-ri-ckov-a, J. Hor-a-ckov-a, and V. Lo-zek, Direct evidence of central European forest refugia during the last glacial period based on mollusc fossils, Quaternary Research, vol.82, pp.222-228, 2014.

P. Kaltenrieder, C. A. Belis, S. Hofstetter, B. Ammann, C. Ravazzi et al., Environmental and climatic conditions at a potential Glacial refugial site of tree species near the Southern Alpine glaciers. New insights from multiproxy sedimentary studies at Lago della Costa (Euganean Hills, Northeastern Italy), Quaternary Sciences Reviews, vol.28, pp.2647-2662, 2009.

W. Kilian, Comptes Rendus de l'Acad emie des Sciences, vol.9, pp.574-576, 1894.

L. Kullman, Early postglacial appearance of tree species in northern Scandinavia: Review and perspective, Quaternary Science Reviews, vol.27, pp.2467-2472, 2008.

G. De-lafontaine, C. A. Guerra, A. Ducousso, and R. J. Petit, Cryptic no more: Soil macrofossils uncover Pleistocene forest microrefugia within a periglacial desert, New Phytologist, vol.204, pp.715-729, 2014.

G. Lang and K. Tobolski, Hobschensee-Late-glacial and Holocene environment of a lake near the timberline, Dissertation Botanicae, vol.87, pp.209-228, 1985.

S. Lavorel, K. Grigulis, G. Leitinger, M. Kohler, U. Schirpke et al., Historical trajectories in land use pattern and grassland ecosystem services in two European alpine landscapes, Regional Environmental Research, vol.17, pp.2251-2264, 2017.

B. Leys, C. Carcaillet, O. Blarquez, A. Lami, S. Musazzi et al., Resistance of mixed subalpine forest to fire frequency changes: The ecological function of dwarf pine (Pinus mugo ssp. mugo), Quaternary Science Reviews, vol.90, pp.60-68, 2014.

K. R. Ludwig and J. B. Paces, Uranium-series dating of pedogenic silica and carbonate, Geochimistry and Cosmochimica Acta, vol.66, pp.487-506, 2002.

C. Miramont, O. Sivan, F. Guibal, B. Kromer, S. Talamo et al., Contribution of French sub-fossil tree-ring chronologies at Barbier River (southern French Alps), vol.22, pp.261-271, 2011.

C. Miramont, O. Sivan, T. Rosique, J. L. Edouard, and M. Jorda, Environmental changes from Allerød to Atlantic, Radiocarbon, vol.42, pp.423-435, 2000.

E. Monnin, A. Inderm?-uhle, A. D?-allenbach, J. Fl?-uckiger, B. Stauffer et al., Atmospheric CO 2 concentrations over the last glacial termination, Science, vol.291, pp.112-114, 2001.

I. H. Myers-smith, S. C. Elmendorf, P. S. Beck, M. Wilmking, M. Hallinger et al., Climate sensitivity of shrub growth across the tundra biome, Nature Climatic Change, vol.5, pp.887-891, 2015.

A. Niskanen, M. Luoto, H. V?-are, and R. K. Heikkinen, Models of Arctic-alpine refugia highlight importance of climate and local topography, Polar Biology, vol.40, pp.489-502, 2017.

L. Öberg and L. Kullman, Ancient subalpine clonal spruces (Picea abies): Sources of postglacial vegetation history in the Swedish Scandes, Arctic, vol.64, pp.183-196, 2011.

E. Ortu, O. Peyron, A. Bordon, J. L. De-beaulieu, C. Siniscalco et al., Lateglacial and Holocene climate oscillations in the South-western Alps: An attempt at quantitative reconstruction. Quaternary International, vol.190, pp.71-88, 2008.
URL : https://hal.archives-ouvertes.fr/halsde-00403091

L. Parducci, K. D. Bennett, G. F. Ficetola, I. G. Alsos, Y. Suyama et al., Ancient plant DNA in lake sediments, New Phytologist, vol.214, pp.924-942, 2017.

L. Parducci, T. Jørgensen, M. M. Tollefsrud, E. Elverland, T. Alm et al., Glacial survival of boreal trees in northern Scandinavia, Science, vol.335, pp.1083-1086, 2012.

A. Pentecost, , 2005.

R. J. Petit, S. Brewer, S. Bordacs, K. Burg, R. Cheddadi et al.,

A. Kremer, Identification of refugia and post-glacial colonisation routes of European white oaks based on chloroplast DNA and fossil pollen evidence, Forest Ecology and Management, vol.156, pp.49-74, 2002.

M. C. Quinzin, S. Normand, S. Dellicour, J. C. Svenning, and P. Mardulyn, Glacial survival of trophically linked boreal species in northern Europe, Proceedings of the Royal Society B, vol.284, 2017.

C. Ravazzi, F. Badino, D. Marsetti, G. Patera, and P. J. Reimer, Glacial to paraglacial history and forest recovery in the Oglio glacier system (Italian Alps) between 26 and 15 ka cal BP, Quaternary Science Reviews, vol.58, pp.146-161, 2012.
DOI : 10.1016/j.quascirev.2012.10.017

C. Ravazzi, M. Donegana, E. Vescovi, E. Arpenti, M. Caccianiga et al., A new Lateglacial site with Picea abies in the northern Apennine foothills: An exception to the model of glacial refugia of trees. Vegetation History and Archaeobotany, vol.15, pp.357-371, 2006.

W. Tinner and P. Kaltenrieder, Rapid responses of high-mountain vegetation to early Holocene environmental changes in the Swiss Alps, Journal of Ecology, vol.93, pp.936-947, 2005.

P. C. Tzedakis, B. C. Emerson, and G. M. Hewitt, Cryptic or mystic? Glacial tree refugia in northern Europe, Trends in Ecology & Evolution, vol.28, pp.696-704, 2013.
DOI : 10.1016/j.tree.2013.09.001
URL : https://doi.org/10.1016/j.tree.2013.09.001

A. Vald-es, J. Lenoir, E. Gallet-moron, E. Andrieu, J. Brunet et al., The contribution of patch-scale conditions is greater than that of macroclimate in explaining local plant diversity in fragmented forests across, Europe. Global Ecology and Biogeography, vol.24, pp.1094-1105, 2015.

M. Walker, S. Johnsen, S. O. Rasmussen, T. Popp, J. P. Steffensen et al., Formal definition and dating of the GSSP (Global Stratotype Section and Point) for the base of the Holocene using the Greenland NGRIP ice core, and selected auxiliary records, Journal of Quaternary Science, vol.24, pp.3-17, 2009.

F. W. Welter-schultes, European non-marine molluscs: A guide for species identification, G? ottingen: Planet Poster Editions, 2012.